adj. (1) Having fruit or cones that remain on the plant without opening for one or more years or (2) having a phenological habit that occurs relatively late in the season (Helms 1998).
(1) Producing fruit or cones that remain on a plant without opening for one or more years. In some plants (some lodgepole pines, for example), cones open and seeds are shed when heat is provided by fire or hot, dry conditions. (2) Having a habit of flowering or fruiting, opening buds, or shedding leaves late in the season relative to other plants in the same habitat – not necessarily related to fire (Helms 1998).